The Making of Mandrill

At the time of his debut in Shanna The She-Devil #4 (June 1973), the villain Mandrill worked with an accomplice named Professor Skecher. Mandrill had the superhuman power to compel women to do his bidding, and the heinous Professor Skecher would tattoo Mandrill's facial markings onto the face of each follower. Sketcher himself had no visible tattoos. The adventurer Shanna O'Hara surprised both men with her athleticism and her ability to resist Mandrill's influence.

By the time the Defenders faced Mandrill, Professor Skecher was out of the picture and Mandrill's new female followers (the Fem-Force) were not tattooed in his image.